Message 2000-09-0010: New critics

Sat, 30 Sep 2000 13:17:36 +0200

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Date: Sat, 30 Sep 2000 13:17:36 +0200
From: David Marjanovic <david.marjanovic@gmx.at>
To: PhyloCode@ouvaxa.cats.ohiou.edu
Subject: New critics

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There's a recent criticism of cladistics and the PhyloCode by Michael J. =
Benton at http://palaeo.gly.bris.ac.uk/phylocode/biolrev.html which has =
just been posted to the Dinosaur Mailing List. Unfortunately the figures =
are not available. Most criticisms there are unfair or point out the =
problems of Sereno's practice of defining (and redefining) taxa so that =
they only make any sense under his present cladogram. However, one =
should have read it, and I strongly suggest to include recommendations =
into the PhyloCode:
  a.. to be very careful with definitions (to take all proposed =
phylogenies into account)
  b.. to include a rather large number of specifiers. For example, I =
would define Aves and/or Avialae something like Neornithes > Oviraptor, =
Troodon, Tyrannosaurus, Deinonychus; in my soon-to-be-published =
phylogeny (see below) oviraptorosaurs are the sister group of =
Metornithes, the group including the usual birds, and if I applicate the =
current definitions of Aves (Archaeopteryx + Neornithes) and Avialae =
(Neornithes > Velociraptor), then Tyrannosaurus, Ornithomimus, Troodon =
and many others belong to both, in other words, they're birds.
  c.. rather not to define a name as long as there is no good consensus =
on its utility
Furthermore, genera (as the last rank) should be kept for quite some =
time after the first version of the PhyloCode becomes active -- =
otherwise, confusion will be endless. The system Sapiens Homo (Hominini =
Homininae Hominidae...) might be an idea for the future (some decades =
from now), but why not turning it by 180=B0 to arrive at (... Hominidae =
Homininae Hominini) Homo sapiens, which is nearly where we are and thus =
would provide stability?
Another approach could be to give every species a "genus" uninomen, but =
whereas this is feasible for nonavian dinosaurs where most genera are =
monotypic, it would hardly be for extant ones like Crocodylus (by the =
way, should we correct that nonsensical y?), Varanus or Bos, not to =
mention insects...

Most genera should not be defined so that stability isn't completely =
destroyed. Archaeopteryx has been defined Archaeopteryx lithographica > =
Neornithes -- in my phylogeny, this includes all dromaeosaurs =
(Deinonychus, Velociraptor...).

Another problematic question is what to do with the standardized endings =
-idae, -inae and -ini (and the many more in ICBN) -- whether to treat =
these endings as random letter combinations that are part of the name or =
whether (as current usage in "dinosaurology") an -idae can't include =
another -idae, and if such turns out to be the case, the less inclusive =
taxon should be renamed into -inae.

Reference:
David Marjanovic: Is Archaeopteryx really a bird? The Dinosaur Society =
Quarterly Review, in press

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<DIV><FONT face=3DArial><FONT size=3D2>There's a recent criticism of =
cladistics and=20
the PhyloCode by Michael J. Benton&nbsp;</FONT><FONT size=3D2>at =
</FONT></FONT><A=20
href=3D"http://palaeo.gly.bris.ac.uk/phylocode/biolrev.html"><FONT =
face=3DArial=20
size=3D2>http://palaeo.gly.bris.ac.uk/phylocode/biolrev.html</FONT></A><F=
ONT=20
face=3DArial size=3D2>&nbsp;which has just been posted&nbsp;to the =
Dinosaur=20
Mailing&nbsp;List.&nbsp;Unfortunately the figures are not available. =
Most=20
criticisms there are unfair or point out the problems of Sereno's =
practice of=20
defining (and redefining) taxa so that they only make any sense under =
his=20
present cladogram. However, one should have read it, and&nbsp;I strongly =
suggest=20
to include recommendations into the PhyloCode:</FONT></DIV>
<UL>
  <LI><FONT face=3DArial size=3D2>to be very careful with definitions =
(to take all=20
  proposed phylogenies into account)</FONT></LI>
  <LI><FONT face=3DArial size=3D2>to include a rather large number=20
  of&nbsp;specifiers. For example, I would define Aves and/or Avialae =
something=20
  like&nbsp;Neornithes &gt; <EM>Oviraptor, Troodon, Tyrannosaurus,=20
  Deinonychus</EM>; in my soon-to-be-published phylogeny (see=20
  below)&nbsp;oviraptorosaurs are the sister group of Metornithes, the =
group=20
  including the usual birds, and if I applicate the current definitions =
of Aves=20
  <EM>(Archaeopteryx</EM> + Neornithes) and Avialae (Neornithes &gt;=20
  <EM>Velociraptor</EM>), then <EM>Tyrannosaurus</EM>, <EM>Ornithomimus, =

  Troodon</EM>&nbsp;and many others belong to both, in other words, =
they're=20
  birds.</FONT></LI>
  <LI><FONT face=3DArial size=3D2>rather not to define a name as long as =
there is no=20
  good consensus on its utility</FONT></LI></UL>
<DIV><FONT face=3DArial size=3D2>Furthermore, genera (as the last rank) =
should be=20
kept for quite some time after the first version of the PhyloCode =
becomes active=20
-- otherwise, confusion will be <STRONG><U>endless</U></STRONG>. The =
system=20
<EM>Sapiens Homo</EM> (<EM>Hominini Homininae Hominidae</EM>...) might =
be an=20
idea for the future (some decades from now), but why not turning it by =
180=B0 to=20
arrive at (... <EM>Hominidae</EM> <EM>Homininae Hominini</EM>) <EM>Homo=20
sapiens</EM>, which is nearly where we are and =
thus&nbsp;would&nbsp;provide=20
stability?</FONT></DIV>
<DIV><FONT face=3DArial size=3D2>Another approach could be to give every =
species a=20
"genus" uninomen, but whereas this is feasible for nonavian dinosaurs =
where most=20
genera are monotypic, it would hardly be for extant ones like=20
<EM>Crocodylus</EM> (by the way, should we correct that nonsensical y?), =

<EM>Varanus</EM> or <EM>Bos</EM>, not to mention insects...</FONT></DIV>
<DIV><FONT face=3DArial size=3D2></FONT>&nbsp;</DIV>
<DIV><FONT face=3DArial size=3D2>Most genera should <STRONG>not be=20
defined&nbsp;</STRONG>so that stability isn't completely=20
destroyed.&nbsp;<EM>Archaeopteryx</EM> has been defined =
<EM>Archaeopteryx</EM>=20
<EM>lithographica</EM> &gt; Neornithes -- in my phylogeny, this includes =
all=20
dromaeosaurs (<EM>Deinonychus</EM>, =
<EM>Velociraptor</EM>...).</FONT></DIV>
<DIV><FONT face=3DArial size=3D2></FONT>&nbsp;</DIV>
<DIV><FONT face=3DArial size=3D2>Another problematic question is what to =
do with the=20
standardized endings -idae, -inae and -ini (and the many more in ICBN) =
--=20
whether to treat these endings as random letter combinations that are =
part of=20
the name or whether (as current usage in "dinosaurology") an -idae can't =
include=20
another -idae, and if such turns out to be the case, the less inclusive =
taxon=20
should be renamed into -inae.</FONT></DIV>
<DIV><FONT face=3DArial size=3D2></FONT>&nbsp;</DIV>
<DIV><FONT face=3DArial size=3D2>Reference:</FONT></DIV>
<DIV><FONT face=3DArial size=3D2>David Marjanovi<FONT =
face=3DArial>&#263;: <STRONG>Is=20
<EM>Archaeopteryx</EM> really a bird?</STRONG> The Dinosaur Society =
Quarterly=20
Review, in press</FONT></FONT></DIV></BODY></HTML>

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